While the link was active, Natani (linktani?) was undoubtably trans- literally a soul being healed over a male pattern, in a female body. Attraction to males isnt counterevidence here- transfolk can still be gay, straight, bi , pan, whatever.
Pre-link youngtani is more of an open question. Strong cultural roles told "her" that her sex meant she was weak. But our first inkling of Natani wanting to "not be female" was shortly after their parents died and Zen blamed Natani's emotional responce (to the death of loved ones!) on femaleness. "I wish I was like you, brother."
There was also a hefty element of not having a female role model, of course. They were parentless, and noone actually cared about youngtani or post-link Natani except Zen, until Keith showed up. Very few even found out about "her":
So culturally, Youngtani was raised as male and was actively suppressing "her" female side... and "Secretly liked it," according to her own soul. If not trans, Youngtani was at least genderfluid.
Post-Link Newtani is an unknown element, of course, though we can take some hints from when Natani was awake while the link was suppressed. Feels pretty action-hero-male-stereotype to me.
But all we've seen so far of Newtani after the link is broken completely is eyes representing a complete soul. If my above speculation on youngtani is correct, I would think that genderfluid youngtani is slotting into a zen-shaped hole in Linktani's psyche, with new experiences and role models that youngtani never had. It will be uncomfortable for a time until Newtani adjusts, but I suspect Newtani will remain male-expressing by wolf standards, but genderfluid at least as far as the Bastinian King's expression of feminine ideal.
